Subgenus (Octopus) J. B. P. A. de Lamarck, 1798

(4-3-2-2-5-2-3-20-8-0-0-0-0)

alecto S. S. Berry, 1953
(4-3-2-2-5-2-3-20-8-1-0-0-0)
Ref. Abbott, 1974 p 594
Keen, 1971 p 894
Nesis, 1987 p 313

americanus H. M. D. de Blainville
(4-3-2-2-5-2-3-20-8-2-0-0-0)
[=Octopus vulgaris G. L. Cuvier, 1797]
Ref. Abbott, 1974 p 593

apollyon (S. S. Berry, 1913)
(4-3-2-2-5-2-3-20-8-3-0-0-0)
[=Octopus dolfleini (Wulker, 1910)]
Ref. Nesis, 1987 p 312

araneoides (Taki, 1964)
(4-3-2-2-5-2-3-20-8-4-0-0-0)
Ref. Nesis, 1987 p 313 d

areolatus deHaan in J. B. Ferussac & A. d'Orbigny, 1840
(4-3-2-2-5-2-3-20-8-5-0-0-0)
[=Octopus ocellatus J. E. Gray, 1849]
Ref. Nesis, 1987 p 309

australis W. E. Hoyle, 1885
(4-3-2-2-5-2-3-20-8-6-0-0-0)
Ref. Nesis, 1987 p 308

bairdii A. E. Verrill, 1873
(4-3-2-2-5-2-3-20-8-7-0-0-0)
Ref. Abbott, 1974 p 594 d

bermudensis W. E. Hoyle, 1885
(4-3-2-2-5-2-3-20-8-8-0-0-0)
[=Octopus macropus J. A. Risso, 1826]
Ref. Abbott, 1974 p 593
Nesis, 1987 p 304

bimaculatus A. E. Verrill, 1883
(4-3-2-2-5-2-3-20-8-9-0-0-0)
California Two-spot Octopus, Two-spotted Octopus
Ref. Abbott, 1974 p 594
Amer. Fish. Soc., 1980 p 152
Keen, 1971 p 894
Nesis, 1987 p 313

bimaculoides G. E. Pickford & B. H. McConnaughey, 1949
(4-3-2-2-5-2-3-20-8-10-0-0-0)
California Two-spot Octopus, Mud-flat Octopus
Ref. Abbott, 1974 p 594
Amer. Fish. Soc., 1980 p 152
Nesis, 1987 p 313 d

briareus G. C. Robson, 1929
(4-3-2-2-5-2-3-20-8-11-0-0-0)
Briar Octopus, Caribbean Reef Octopus, Reef Octopus
Ref. Abbott, 1974 p 593 d
Abbott & Dance, 1982 p 377 c
Amer. Fish. Soc., 1980 p 152
Nesis, 1987 p 304 c d
Voss, Opresko & Thomas, 1973 p 12 bw

brocki A. E. Ortmann, 1888
(4-3-2-2-5-2-3-20-8-12-0-0-0)
[=Octopus mambranaceus J. R. C. Quoy & J. P. Gaimard, 1832]
Ref. Nesis, 1987 p 309

brucei (W. E. Hoyle, 1912)
(4-3-2-2-5-2-3-20-8-13-0-0-0)
[=Octopus megalocyathus A. A. Gould, 1846]
Ref. Nesis, 1987 p 305

burryi G. L. Voss, 1950[1953]
(4-3-2-2-5-2-3-20-8-14-0-0-0)
Brownstripe Octopus, Burry's Octopus
Ref. Abbott, 1974 p 593 d
Amer. Fish. Soc., 1980 p 152
Syn. Octopus vincenti G. E. Pickford, 1955
Ref. Nesis, 1987 p 304 d

californicus (S. S. Berry, 1911)
(4-3-2-2-5-2-3-20-8-15-0-0-0)
Orange Bigeye Octopus
Ref. Abbott, 1974 p 594
Amer. Fish. Soc., 1980 p 152
Nesis, 1987 p 313

carolinensis A. E. Verrill,
(4-3-2-2-5-2-3-20-8-16-0-0-0)
[=Octopus vulgaris G. L. Cuvier, 1797]
Ref. Abbott, 1974 p 593

chierchiae G. Jatta, 1889
(4-3-2-2-5-2-3-20-8-17-0-0-0)
Ref. Keen, 1971 p 894
Nesis, 1987 p 314

chromatus L. G. Heilprin, 1888
(4-3-2-2-5-2-3-20-8-18-0-0-0)
[=Octopus macropus J. A. Risso, 1826]
Ref. Abbott, 1974 p 593

conispadiceus (M. Sasaki, 1917)
(4-3-2-2-5-2-3-20-8-19-0-0-0)
Ref. Nesis, 1987 p 309 d

cyaneus (4-3-2-2-5-2-3-20-8-20-0-0-0)
Ref. Nesis, 1987 p 71 c

defilipppi Verany, 1851
(4-3-2-2-5-2-3-20-8-21-0-0-0)
Longarm Octopus
Ref. Amer. Fish. Soc., 1980 p 152
Nesis, 1987 p 304 d

digueti E. Perrier & A. T. de Rochebrune, 1894
(4-3-2-2-5-2-3-20-8-22-0-0-0)
Pacific Pygmy Octopus
Ref. Amer. Fish. Soc., 1980 p 152
Keen, 1971 p 894
Nesis, 1987 p 314

dolfleini (Wulker, 1910)
(4-3-2-2-5-2-3-20-8-23-0-0-0)
Common Pacific Octopus, Giant Octopus
Ref. Amer. Fish. Soc., 1980 p 152
Kozloff, 1987 p 295 d
Syn. Octopus hongkongensis W. E. Hoyle;
Octopus punctatus W. M. Gabb, 1862
Ref. Abbott, 1974 p 593
Syn. Octopus apollyon (S. S. Berry, 1913);
Octopus gilbertianus (S. S. Berry, 1912);
Octopus hongkongensis authors [not W. E. Hoyle];
? Octopus madokai (S. S. Berry, 1921);
Octopus punctatus authors [not H. M. D. de Blainville, 1826;
paroctopus asper Akimushkin, 1963
Ref. Nesis, 1987 p 312 d

apollyon S. S. Berry, 1912[1913]
(4-3-2-2-5-2-3-20-8-23-1-0-0)
Syn. Octopus gilbertianus S. S. Berry, 1912
Ref. Abbott, 1974 p 594
Nesis, 1987 p 312 d

dofleini Wulker, 1910
(4-3-2-2-5-2-3-20-8-23-2-0-0)
Ref. Nesis, 1987 p 312 c
Syn. Octopus madokai S. S. Berry, 1921;
Octopus pustulosus M. Sasaki, 1920
[not H. M. D. de Blainville, 1826]
Ref. Abbott, 1974 p 594

martini G. E. Pickford, 1964
(4-3-2-2-5-2-3-20-8-23-3-0-0)
Ref. Abbott, 1974 p 594
Nesis, 1987 p 312 d

fangsiao A. d'Orbigny, 1840
(4-3-2-2-5-2-3-20-8-24-0-0-0)
[=Octopus ocellatus J. E. Gray, 1849]
Ref. Nesis, 1987 p 309

fasciata W. E. Hoyle, 1886
(4-3-2-2-5-2-3-20-8-25-0-0-0)
[=Hapalochlaena maculosa (W. E. Hoyle, 1883)]
Ref. Nesis, 1987 p 293

fitchi S. S. Berry, 1953
(4-3-2-2-5-2-3-20-8-26-0-0-0)
Lilliput Octopus
Ref. Abbott, 1974 p 594
Amer. Fish. Soc., 1980 p 152
Keen, 1971 p 894
Nesis, 1987 p 314

fontanianus A. d'Orbigny, 1835
(4-3-2-2-5-2-3-20-8-27-0-0-0)
Syn. Joubinia fontaniana authors;
Robsonella fontaniana authors
Ref. Nesis, 1987 p 308

Var. africanus (G. C. Robson, 1929)
(4-3-2-2-5-2-3-20-8-27-0-0-1)
Ref. Nesis, 1987 p 308

fujitai (M. Sasaki, 1929)
(4-3-2-2-5-2-3-20-8-28-0-0-0)
Ref. Nesis, 1987 p 312 d

gilbertianus S. S. Berry, 1912
(4-3-2-2-5-2-3-20-8-29-0-0-0)
[=Octopus dolfleini apollyon S. S. Berry, 1912[1913]]
Ref. Abbott, 1974 p 594
Nesis, 1987 p 312
[=Octopus dolfleini (Wulker, 1910)]
Ref. Nesis, 1987 p 312

hongkongensis authors [not W. E. Hoyle]
(4-3-2-2-5-2-3-20-8-30-0-0-0)
[=Octopus dolfleini (Wulker, 1910)]
Ref. Nesis, 1987 p 312

hongkongensis W. E. Hoyle
(4-3-2-2-5-2-3-20-8-31-0-0-0)
[=Octopus dolfleini (Wulker, 1910)]
Ref. Abbott, 1974 p 593

hubbsorum S. S. Berry, 1953
(4-3-2-2-5-2-3-20-8-32-0-0-0)
Ref. Abbott, 1974 p 594
Keen, 1971 p 894
Nesis, 1987 p 314

hummelincki W. Adam, 1936
(4-3-2-2-5-2-3-20-8-33-0-0-0)
Caribbean Two-spot Octopus, Seedweed Octopus
Ref. Abbott, 1974 p 593
Amer. Fish. Soc., 1980 p 152
Nesis, 1987 p 303 d

hyadesi A. T. de Rochebrune & J. Mabille, 1887
(4-3-2-2-5-2-3-20-8-34-0-0-0)
Ref. Nesis, 1987 p 305

joubini G. C. Robson, 1929
(4-3-2-2-5-2-3-20-8-35-0-0-0)
Atlantic Pygmy Octopus, Joubin's Octopus, Pygmy Octopus
Ref. Amer. Fish. Soc., 1980 p 152
Voss, Opresko & Thomas, 1973 p 16 bw
Syn. Octopus mercatoris W. Adam, 1937;
Paroctopus joubini G. C. Robson, 1929
Ref. Abbott, 1974 p 593 bw d
Nesis, 1987 p 305 d

leioderma (S. S. Berry, 1911)
(4-3-2-2-5-2-3-20-8-36-0-0-0)
Smoothskin Octopus
Ref. Abbott, 1974 p 594
Amer. Fish. Soc., 1980 p 152
Kozloff, 1987 p 295
Nesis, 1987 p 309,313 d

lobensis Z. J. A. de Castellanos & Menni, 1969
(4-3-2-2-5-2-3-20-8-37-0-0-0)
Ref. Nesis, 1987 p 308 d

longispadiceus (M. Sasaki, 1917)
(4-3-2-2-5-2-3-20-8-38-0-0-0)
Ref. Nesis, 1987 p 312 d

macropus authors [not J. A. Risso, 1826]
(4-3-2-2-5-2-3-20-8-39-0-0-0)
[=Octopus variabilis (M. Sasaki, 1929)]
Ref. Nesis, 1987 p 309

Var. minor (M. Sasaki, 1920)
(4-3-2-2-5-2-3-20-8-39-0-0-1)
[=Octopus variabilis (M. Sasaki, 1929)]
Ref. Nesis, 1987 p 309

macropus J. A. Risso, 1826
(4-3-2-2-5-2-3-20-8-40-0-0-0)
Grass Octopus, White Spotted Octopus, White-spotted Octopus
Ref. Amer. Fish. Soc., 1980 p 152
Voss, Opresko & Thomas, 1973 p 10 bw
Syn. Octopus bermudensis W. E. Hoyle, 1885;
Octopus chromatus L. G. Heilprin, 1888
Ref. Abbott, 1974 p 593 d
Nesis, 1987 p 304 c d

madokai S. S. Berry, 1921
(4-3-2-2-5-2-3-20-8-41-0-0-0)
[=Octopus dofleini dofleini (Wulker, 1910)]
Ref. Abbott, 1974 p 594
[?=Octopus dofleini (Wulker, 1910)]
Ref. Nesis, 1987 p 312

maya Voss & Solis, 1966
(4-3-2-2-5-2-3-20-8-42-0-0-0)
Mexican Four-eyed Octopus
Ref. Nesis, 1987 p 303 d
Voss, Opresko & Thomas, 1973 p 8 bw

megalocyathus A. A. Gould, 1846
(4-3-2-2-5-2-3-20-8-43-0-0-0)
Syn. Octopus brucei (W. E. Hoyle, 1912);
Octopus patagonicus E. Lonnberg, 1898
Ref. Nesis, 1987 p 305

membranaceus J. R. C. Quoy & J. P. Gaimard, 1832
(4-3-2-2-5-2-3-20-8-44-0-0-0)
Syn. Octopus brocki A. E. Ortmann, 1888;
Octopus ovulum (M. Sasaki, 1917);
Octopus pulcher Broch, 1887
Ref. Nesis, 1987 p 309

membranaceus A. T. de Rochebrune & J. Mabille, 1887 [not J. R. C. Quoy & J. P. Gaimard, 1832]
(4-3-2-2-5-2-3-20-8-45-0-0-0)
Ref. Nesis, 1987 p 305 d

mercatoris W. Adam, 1937
(4-3-2-2-5-2-3-20-8-46-0-0-0)
[=Octopus joubini G. C. Robson, 1929]
Ref. Abbott, 1974 p 593
Nesis, 1987 p 305

micropyrsus S. S. Berry, 1953
(4-3-2-2-5-2-3-20-8-47-0-0-0)
Ref. Abbott, 1974 p 594
Nesis, 1987 p 314

occidentalis W. E. Hoyle, 1896
(4-3-2-2-5-2-3-20-8-48-0-0-0)
Ref. Keen, 1971 p 894

ocellatus J. E. Gray, 1849
(4-3-2-2-5-2-3-20-8-49-0-0-0)
Syn. Octopus areolatus deHaan in J. B. Ferussac & A. d'Orbigny, 1840;
Octopus fangsiao A. d'Orbigny, 1840
Ref. Nesis, 1987 p 309 c

ochotensis (M. Sasaki, 1920)
(4-3-2-2-5-2-3-20-8-50-0-0-0)
Ref. Nesis, 1987 p 309

oculifer (W. E. Hoyle, 1904)
(4-3-2-2-5-2-3-20-8-51-0-0-0)
Ref. Keen, 1971 p 894

ornatus
(4-3-2-2-5-2-3-20-8-52-0-0-0)
Ref. Nesis, 1987 p 29 c

oshimai (M. Sasaki, 1929)
(4-3-2-2-5-2-3-20-8-53-0-0-0)
Ref. Nesis, 1987 p 315

ovulum (M. Sasaki, 1917)
(4-3-2-2-5-2-3-20-8-54-0-0-0)
[=Octopus membranaceus J. R. C. Quoy & J. P. Gaimmard, 1832]
Ref. Nesis, 1987 p 309

patagonicus E. Lonnberg, 1898
(4-3-2-2-5-2-3-20-8-55-0-0-0)
[=Octopus megalocyathus A. A. Gould, 1846]
Ref. Nesis, 1987 p 305

penicillifer S. S. Berry, 1954
(4-3-2-2-5-2-3-20-8-56-0-0-0)
Ref. Abbott, 1974 p 594
Keen, 1971 p 894
Nesis, 1987 p 315

pentherinus A. T. de Rochebrune & J. Mabille, 1887
(4-3-2-2-5-2-3-20-8-57-0-0-0)
Ref. Nesis, 1987 p 305

pictus Brock, 1882 [not H. M. D. de Blainville]
(4-3-2-2-5-2-3-20-8-58-0-0-0)
[=Hapalochlaena maculosa (W. E. Hoyle, 1883)]
Ref. Nesis, 1987 p 293

? pricei S. S. Berry, 1913
(4-3-2-2-5-2-3-20-8-59-0-0-0)
Ref. Abbott, 1974 p 594
Nesis, 1987 p 315

pulcher Broch, 1887
(4-3-2-2-5-2-3-20-8-60-0-0-0)
[=Octopus membranaceus J. R. C. Quoy & J. P. Gaimard, 1832]
Ref. Nesis, 1987 p 309

punctatus authors [not H. M. D. de Blainville, 1826]
(4-3-2-2-5-2-3-20-8-61-0-0-0)
[=Octopus dofleini (Wulker, 1910)]
Ref. Nesis, 1987 p 312

punctatus W. M. Gabb, 1862 [not H. M. D. de Blainville, 1826
(4-3-2-2-5-2-3-20-8-62-0-0-0)
[=Octopus dofleini (Wulker, 1910)]
Ref. Abbott, 1974 p 594

pustulosus M. Sasaki, 1920 [not H. M. D. de Blainville, 1826]
(4-3-2-2-5-2-3-20-8-63-0-0-0)
[=Octopus dofleini dofleini (Wulker, 1910)]
Ref. Abbott, 1974 p 594

roosevelti H. G. Stuart, 1941
(4-3-2-2-5-2-3-20-8-64-0-0-0)
Ref. Keen, 1971 p 894

rubescens S. S. Berry, 1953
(4-3-2-2-5-2-3-20-8-65-0-0-0)
Red Octopus
Ref. Abbott, 1974 p 594
Amer. Fish. Soc., 1980 p 152
Kozloff, 1987 p 295 d
Nesis, 1987 p 314

rugosus authors [not L. A. G. Bosc, 1792]
(4-3-2-2-5-2-3-20-8-66-0-0-0)
[=Octopus vulgaris G. L. Cuvier, 1797]
Ref. Abbott, 1974 p 593

rugosus (L. A. G. Bosc, 1792)
(4-3-2-2-5-2-3-20-8-67-0-0-0)
[=Octopus vulgaris J. B. P. A. de Lamarck, 1798]
Ref. Nesis, 1987 p 304

salutii Verany, 1851
(4-3-2-2-5-2-3-20-8-68-0-0-0)
Ref. Nesis, 1987 p 305 d

schultzei (W. E. Hoyle, 1910)
(4-3-2-2-5-2-3-20-8-69-0-0-0)
Ref. Nesis, 1987 p 304

scorpio S. S. Berry, 1920
(4-3-2-2-5-2-3-20-8-70-0-0-0)
[=Scaeurgus unicirrhus (A. d'Orbigny, 1840)]
Ref. Abbott, 1974 p 595

spinosus (M. Sasaki, 1920)
(4-3-2-2-5-2-3-20-8-71-0-0-0)
Ref. Nesis, 1987 p 313

stictochrus G. L. Voss, 1971
(4-3-2-2-5-2-3-20-8-72-0-0-0)
Ref. Nesis, 1987 p 315

tehuelchus A. d'Orbigny, 1835
(4-3-2-2-5-2-3-20-8-73-0-0-0)
Ref. Nesis, 1987 p 308 d

tsugarensis (M. Sasaki, 1920)
(4-3-2-2-5-2-3-20-8-74-0-0-0)
Ref. Nesis, 1987 p 312 d

variabilis (M. Sasaki, 1929)
(4-3-2-2-5-2-3-20-8-75-0-0-0)
Syn. Octopus macroppus authors [not J. A. Risso, 1826];
Octopus macropus Var. minor (M. Sasaki, 1920);
Octopus variabilis Var. minor (M. Sasaki, 1929)
Ref. Nesis, 1987 p 309 c d

Var. minor (M. Sasaki, 1929)
(4-3-2-2-5-2-3-20-8-75-0-0-1)
[=Octopus variabilis (M. Sasaki, 1929)]
Ref. Nesis, 1987 p 309

veligero S. S. Berry, 1953
(4-3-2-2-5-2-3-20-8-76-0-0-0)
Ref. Abbott, 1974 p 594
Nesis, 1987 p 314

vincenti G. E. Pickford, 1955
(4-3-2-2-5-2-3-20-8-77-0-0-0)
[=Octopus burryi G. L. Voss, 1950]
Ref. Nesis, 1987 p 304

vulgaris J. B. P. A. de Lamarck, 1798 [T]
(4-3-2-2-5-2-3-20-8-78-0-0-0)
Common Octopus
Ref. Abbott, 1974 p 592
Amer. Fish. Soc., 1980 p 152
Syn. Octopus rugosus (L. A. G. Bosc, 1792)
Ref. Nesis, 1987 p 292,304 c d

vulgaris G. L. Cuvier, 1797
(4-3-2-2-5-2-3-20-8-79-0-0-0)
Common Atlantic Octopus, Common Octopus
Ref. Voss, Opresko & Thomas, 1973 p 14 bw
Syn. Octopus americanus H. M. D. de Blainville;
Octopus carolinensis A. E. Verrill;
Octopus rugosus authors
Ref. Abbott, 1974 p 593 d

yendoi (M. Sasaki, 1920)
(4-3-2-2-5-2-3-20-8-80-0-0-0)
Ref. Nesis, 1987 p 312 d

zonatus G. L. Voss, 1966
(4-3-2-2-5-2-3-20-8-81-0-0-0)
Ref. Nesis, 1987 p 303 d